After beating the game, I felt bad that I had killed the one girl. The ending made it quite clear that these girls are human.
I saved them all until the reveal, then I killed them.
MUAHAHAHAH.
How does saving them all relate to Tonic Collector?
That’s one of the achievements I didn’t get.
At least one of the teddies contains a tonic you can’t get any other way.
Two tonics can only be obtained from Tennenbaum by saving the LSs. And you need all 53.